Peering at New York City’s landscape more than 400 years in the past — when the marshes, ponds and streams crisscrossed our string of islands — can help planners and policymakers better understand the city’s flooding future.

Researchers at the New York Botanical Garden looked at where water used to be, where there’s water now and where — thanks to climate change — water will be in the coming years. Those places are called Blue Zones.

The five boroughs contain more than 500 of them, according to a new paper published Wednesday in the Annals of the New York Academy of Sciences, the first comprehensive analysis of its kind. The Blue Zones cover more than one-fifth of New York’s land.